A black crested macaque male with an inscrutable expression on his face. Macaques have about the same number of ‘mimetic muscles’, the muscles that create facial expressions, as humans. Out of all studied macaque species, the black crested macaque has the richest repertoire of facial expressions. Unlike other primates I have worked with, the crested macaque will look you right in the eye, almost seeming to be evaluating you. Tangkoko, Sulawesi.
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